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| The heart and soul of stems are
the vascular bundles. Vascular bundles play a very important
role in the survival of the plants. Without the vascular bundles,
plants would not be able to transport the important materials they need
for survival.
As we have previously discussed, the organization of the monocot and dicot stem differs and we can recognize a stem as being monocot or dicot by examining this organization. Likewise, the organization within these vascular bundles differs and this can be used to identify the bundle as monocot or dicot. First, let's examine the types of tissues found within all vascular bundles. All vascular bundles contain xylem. Xylem is a tissue that conducts water and dissolved minerals from the roots up to the leaves. It is easy to think of xylem as a set of straws connected end to end. Just like a straw, when water (with dissolved minerals) is pulled through the upper part of the xylem, water is pulled through the bottom parts of the xylem due to the surface tension of water. What causes the water to be pulled in the first place? That is a topic for a later lesson, but it involves a process (transpiration) where water is removed from the leaves. The other primary tissue found in vascular bundles is phloem. Phloem is a collection of cells that carry nutrients, in the form of carbohydrates, from the leaves to the roots. Leaves are commonly thought of as the main site of photosynthesis, the process which converts carbon dioxide into usable carbohydrates. It should be fairly obvious that gravity plays a role in the movement of materials through the phloem. |
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As we take a closer look at the vascular bundle of a monocot, we notice a striking resemblance to the face of an alien or a monster (the monocot monster). The charcteristic "eyes" of the monster is the xylem while the "forehead" of the monster is phloem. The xylem is typically oriented towards the inside of the stem while the phloem is typically oriented towards the outside. The entire vascular bundle is surrounded by
a bundle sheath, which is a collection of sclerenchyma cells.
The bundle sheath offers support for the bundle. The surrounding
tissue is ground tissue and is composed of parenchyma cells.
The ground tissue serves two main functions; support and storage.
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The vascular bundle of a monocot stem. |
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The dicot vascular bundle is quite different from that of a monocot. In many dicots, especially herbaceous dicots, the most externally oriented portion of the bundle is the phloem fiber cap. This structure strengthens the stem and sits just "on top" of the phloem. The next predominate layer is the xylem. However, the phloem and the xylem are separated by the vascular cambium. The vascular cambium is a single layer of cells that is a lateral meristem. Remember, a lateral meristem is an area of rapidly dividing cells that is responsible for secondary growth. Secondary growth results in the thickening of plant stems. The meristematic cells found in this region divide into either secondary xylem or phloem. The dicot vascular bundle is surrounded by the cortex near the epidermis and the pith towards the center of the stem. Both are composed of a variety of cells and perform a range of activities much like that of the monocot's pith. |
| Now take the time to pull out a sheet of paper and sketch each of the two vascular bundles, again. This time be sure to include all of the cells and tissue discussed above. Remember to make your drawings neat enough so that you can identify and label each cell and tissue. |
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Now it's time to practice bundle identification. Do this by selecting this site, and following the instructions. Label your paper Stem 1 and sketch the cross section. Then label A, B, C, D, E, F down the left side of your paper. Identify each structure, then label the stem cross section as a monocot or dicot. |
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You should be getting the hang of identifying a monocot and a dicot both by the stem cross section and by the vascular bundle. But there are a much wider range of stems than the tiny samples viewed with the microscope. Obviously a cross section of a giant redwood would not fit on a slide. So how do these stems get so big? That's next on the investigation. Continue to stem thickening.
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